April Jones had a "happy face" when she got into a vehicle shortly before her disappearance, a court has heard.
April's best friend, seen in video evidence at Mold Crown Court, said the five-year-old appeared to go willingly and did not cry.
The court watched a police interview with the seven-year-old recorded the day after April went missing.
The girl, clutching a toy bunny during her testimony, described the vehicle April got into as a grey Land Rover and that she had seen it before.
Speaking about the person in the vehicle, she said: "He got out of the van and was waiting outside the van for someone".
April got in through the "drivers way" and then sat in the back seat, added the seven-year-old, who said "she wasn't crying, she was happy".
The vehicle then drove off, the court heard.
The accused, Mark Bridger, shook his head when the girl described the man with the Land Rover as having green or blue eyes and brown hair, and probably wearing a green jacket.
The young witness also described how April's brother came looking for his sister to take her home and how shortly after the alarm was raised.
Defence lawyers will be able to cross-examine the girl later this afternoon.
Mark Bridger, 47, has denied abducting and murdering the five-year-old, but says he ran April over and does not remember what he did with her body.
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